Franz, Richard, and Irvy Quitmyer. “A Fossil and Zooarchaeological History of the Gopher Tortoise (Gopher Polyphemus) in the Southeastern United States”. Bulletin of the Florida Museum of Natural History 45, no. 4 (December 31, 2005): 179–200. Accessed April 24, 2026. https://flmnhbulletin.com/index.php/flmnh/article/view/flmnh-vol45-no4-pp179-200.